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The Carnian/Norian boundary succession at Berlin-Ichthyosaur State Park (Upper Triassic, central Nevada, USA)

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Abstract

The Upper Carnian-Lower Norian (Upper Triassic) Luning Formation at Berlin-Ichthyosaur State Park (BISP) in central NV (USA) has been sampled using for the first time the bed-by-bed approach for ammonoids, pelagic bivalves, and conodonts, more than 60 years after its first description by Silberling (U.S. Geological Survey Professional Paper 322: 1–63, 1959). BISP is historically important for the definition of the uppermost Carnian of the North American Triassic chronostratigraphic scale and is known worldwide as one of the most important ichthyosaur Fossil-Lagerstätte because of its extraordinary record of 37 articulated, large-sized specimens of Shonisaurus popularis. Nearly 190 ammonoids were collected from two stratigraphic sections, documenting all the latest Carnian to Early Norian ammonoid faunas previously described by Silberling. Halobiids were collected from five levels, and the first report of conodonts from BISP includes faunas from 13 levels. The ~340-m thick Brick Pile section, the most complete in the study area, includes the uppermost Carnian Macrolobatus Zone, which provides conodont faunas of the lower primitia zone and Halobia septentrionalis. The 200-m thick lowermost Norian Kerri Zone, which begins 52 m above the Macrolobatus Zone, yields conodonts of the upper primitia zone in its lower part, together with H. cf. beyrichi and H. cf. selwyni. The ichthyosaur-bearing interval, whose stratigraphic position has been interpreted quite differently by previous authors, is documented in the uppermost Carnian Macrolobatus Zone and is characterized by rich Tropites-dominated ammonoid faunas and by the onset of Halobia. All models proposed by various workers to explain the unusual ichthyosaur record are discussed and an additional explanation for the main ichthyosaur-bearing bed is proposed. The new hypothesis is that a harmful algal bloom (HAB) may have been the trigger for the mass mortality recorded in this level. Although the C/N boundary in the Brick Pile section lies within a 52 m interval that presently lacks paleontologic data, this succession is included in a small group of sections that are expected to contribute to the definition of the GSSP of the Norian stage. Correlation of the Brick Pile section with the best Carnian/Norian sections in northeastern British Columbia is discussed. Compared to the British Columbia Juvavites Cove and the GSSP candidate Black Bear Ridge sections, the Brick Pile section exhibits an ammonoid and Halobia record that is slightly more similar to that of the Tethyan sections. Correlation of the Brick Pile section with the second GSSP candidate Pizzo Mondello (Sicily, Italy) well demonstrates the significant problems encountered in calibration of the Tethyan and North American scales.

Kurzfassung

Die oberkarnische bis unternorische (Späte Trias) Luning Formation im Berlin-Ichthyosaur State Park (BISP) in Zentral-Nevada (USA) wurde mehr als 60 Jahre nach der ersten Beschreibung durch Silberling (U. S. Geological Survey Professional Paper 322: 1–63, 1959) erstmals bankweise auf Ammoniten, pelagische Bivalven und Conodonten untersucht. Der BISP ist von historischer Bedeutung für die Definition des obersten Karn in der chronostratigraphischen Tabelle der nordamerikanischen Trias und ist eine der weltweit bekanntesten Ichthyosaurier Fossillagerstätten, ausgezeichnet durch den außergewöhnlichen Fund von 37 artikulierten, großen Exemplaren von Shonisaurus popularis. Nahezu 190 Ammoniten aus zwei stratigraphischen Abschnitten belegen oberstes Karn bis Unternor, wie schon früher durch Silberling beschrieben. Halobiiden liegen aus fünf Niveaus vor, und der Erstnachweis von Conodonten im BISP beinhaltet Faunen aus 13 Niveaus. Der ca. 340 m mächtige Brick Pile Aufschluss ist der kompletteste im untersuchten Gebiet. Er schließt die rund 20 m mächtige Macrolobatus Zone des obersten Karn mit Conodonten-Faunen aus der unteren primitia zone und Halobia septentrionalis ein. Nach einer Lücke im Fossilbefund von 52 m folgt die 200 m mächtige Kerri Zone des untersten Nor, welche im unteren Teil Conodonten der oberen primitia zone gemeinsam mit H. cf. beyrichi und H. cf. selwyni erbrachte. Das Ichthyosaurier führende Intervall, von bisherigen Autoren teils ins Karn und/oder ins Nor eingestuft, ist durch eine ergiebige Tropites-Fauna und das Einsetzen von Halobia charakterisiert und kann damit in das obere Karn eingestuft werden. Alle Modelle, die von verschiedenen Autoren für die Erklärung des außergewöhnlichen Ichthyosaurier-Vorkommens vorgeschlagen wurden, werden diskutiert und um ein zusätzliches Genesemodell für die Haupt-Ichthyosaurier-Bank erweitert. Nach der neuen Hypothese führte wohl eine schädliche Algenblüte zu einem Massensterben in diesem Horizont. Obwohl die Karn/Nor Grenze des Brick Pile Aufschlusses im Bereich des fossilleeren 52-m-Intervalls liegt, wird er zu der kleinen Gruppe von Profilen gezählt, welche wesentlich zur Definition des GSSP der Norischen Stufe beitragen können. Die Korrelation von Brick Pile mit den wichtigsten Karn/Nor-Grenzprofilen von British Columbia und Sizilien wird diskutiert. Verglichen mit Juvavites Cove und dem GSSP-Kanditaten Black Bear Ridge in B.C., weist Brick Pile bei den Ammonoideen und Halobien einen etwas stärkeren Tethys-Bezug auf. Ein Vergleich von Brick Pile mit dem zweiten GSSP-Kandidaten Pizzo Mondello (Sizilien, Italien) zeigt die signifikanten Probleme, welche bei der Korrelation tethyaler und nordamerikanischer Biochronologien auftreten.

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References

  • Balini, M., and C. Renesto. 2012. Cymbospondylus vertebrae (Ichthyosauria, Shastasauridae) from the Late Anisian Prezzo Limestone (Middle Triassic, Southern Alps) with an overview of the chronostratigraphic distribution of the group. Rivista Italiana di Paleontologia e Stratigrafia 118: 155–172.

    Google Scholar 

  • Balini, M., S.G. Lucas, J.F. Jenks, and J.A. Spielmann. 2010a. Triassic ammonoid biostratigraphy: an overview. In The Triassic Timescale, ed. S.G. Lucas, 221–262. Geological Society, Special Publications 334.

  • Balini, M., A. Bertinelli, P. Di Stefano, C. Guaiumi, M. Levera, M. Mazza, G. Muttoni, A. Nicora, N. Preto, and M. Rigo. 2010b. The Late Carnian-Rhaetian succession at Pizzo Mondello (Sicani Mountains). Albertiana 39: 36–57.

    Google Scholar 

  • Balini, M., L. Krystyn, M. Levera, and A. Tripodo. 2012. Late Carnian-Early Norian ammonoids from the GSSP candidate section Pizzo Mondello (Sicani Moutains, Sicily). Rivista Italiana di Paleontologia e Stratigrafia 118: 47–84.

    Google Scholar 

  • Bargu, S., C.L. Powell, Z. Wang, G.J. Doucette, and M.W. Silver. 2008. Note on the occurrence of Pseudo-nitzschia australis and domoic acid in squid from Monterey Bay, CA (USA). Harmful algae 7: 45–51.

    Article  Google Scholar 

  • Bottjer, D.J. 2002. Berlin-Ichthyosaur: preserving some of the Earth’s largest marine vertebrates. In Exceptional fossil preservation: a unique view on the evolution of marine life, ed. D.J. Bottjer, W. Etter, J.W. Hagadorn, and C.M. Tang, 243–250. New York: Columbia University Press.

    Google Scholar 

  • Bottjer, D.J., W. Etter, J.W. Hagadorn, and C.M. Tang. 2002. Fossil-Lagerstätten: Jewels of the Fossil Record. In Exceptional fossil preservation: a unique view on the evolution of marine life, ed. D.J. Bottjer, W. Etter, J.W. Hagadorn, and C.M. Tang, 1–10. New York: Columbia University Press.

    Google Scholar 

  • Bychkov, Y.M. 1995. Late Triassic trachiceratids and sirenitids from the upper stream of the Yana Okhotskaya River basin, 1–67. Magadan: Severo-Vostocnyj Nauchnyj Center Dalnevostochnogo Otdeleniya RAN. (in Russian).

    Google Scholar 

  • Callaway, J.M., and J.A. Massare. 1989. Geographic and Stratigraphic Distribution of the Triassic Ichthyosauria (Reptilia; Diapsida). Neues Jahrbuch für Geologie und Paläontologie 178: 37–58.

    Google Scholar 

  • Camp, C.L. 1976. Vorläufige Mitteilungen über große Ichthyosaurier aus der oberen Trias von Nevada. Österreichischen Akademie der Wissenschaften, Sitzungsberichte der mathematisch-naturwissenschaftlichen Klasse Abt. I 185: 125–134.

    Google Scholar 

  • Camp, C.L. 1980. Large Ichthyosaurs from the Upper Triassic of Nevada. Palaeontograhica Abt. A 170: 139–200.

    Google Scholar 

  • Campbell, H.J. 1994. The Triassic bivalves Halobia and Daonella in New Zealand, New Caledonia, and Svalbard. Institute of Geological & Nuclear Sciences Monograph 4: 1–166.

    Google Scholar 

  • Carter, E.S., and M.J. Orchard. 2013. Intercalibration of conodont and radiolarian faunas from the Carnian-Norian Boundary Interval in Haida Gwaii, British Columbia, Canada. In The Triassic system, ed. L.H. Tanner, J.A. Spielman, and S.G. Lucas, 67–92. New Mexico Museum of Natural History and Science, Bulletin 61.

  • Costa, P.R., R. Rosa, J. Pereira, and M.A.M. Sampayo. 2005. Detection of domoic acid, the amnesic shellfish toxin, in the digestive gland of Eledone cirrhosa and E. moschata (Cephalopoda, Octopoda) from the Portuguese coast. Aquatic Living Resources 18: 395–400.

    Article  Google Scholar 

  • Dagys, A.S. 1988. Major features of the geographic differentiation of Triassic ammonoids. In Cephalopods present and past, ed. J. Wiedmann, and J. Kullmann, 341–349. Stuttgart: Schweizerbart’sche Verlagsbuchhandlung.

    Google Scholar 

  • De Capoa Bonardi, P. 1984. Halobia zones in the pelagic Late Triassic sequences of the central Mediterranean area. Bollettino della Società Paleontologica Italiana 23: 91–102.

    Google Scholar 

  • Ferguson, H.G. and S.H. Cathcart. 1954. Geology of the Round Mountain quadrangle, Nevada. In U. S. Geological Survey Geological Quadrangle Map 40.

  • Gruber, B. 1976. Neue ergebnisse auf dem gebiete der Ökologie, Stratigraphie und Phylogenie der Halobien (Bivalvia). Mitteilungen der Gesellschaft der Geologie und Bergbaustudenten in Österreich 23: 181–198.

    Google Scholar 

  • Hogler, J.A. 1992. Taphonomy and paleoecology of Shonisaurus popularis (Reptilia: Ichthyosauria). Palaios 7: 108–117.

    Article  Google Scholar 

  • Hogler, J.A., and R.A. Hanger. 1989. A new Chondrophorine (Hydrozoa, Velellidae) from the Upper Triassic of Nevada. Journal of Paleontology 63: 249–251.

    Google Scholar 

  • Kosch, B.F. 1990. A revision of the skeletal reconstruction of Shonisaurus popularis (Reptilia: Ichthyosauria). Journal of Vertebrate Paleontology 10: 512–514.

    Article  Google Scholar 

  • Kristan-Tollmann, E., and A. Tollmann. 1983. Tethys-Faunelemente in der Trias der USA. Mitteilungen der Osterreichischen Geologischen Gesellschaft 76: 213–272.

    Google Scholar 

  • Krystyn, L. 1973. Zur Ammoniten- und Conodonten-Stratigraphie der Hallstätter Obertrias (Salzkammergut, Österreich). Verhandlungen der Geologischen Bundesanstalt, 1973/1:113–153.

  • Krystyn, L. 1974. Zur Grenzziehung Karn-Nor mit Ammoniten und Conodonten. Österreichischen Akademie der Wissenschaften, Anzeiger der mathematisch-naturwissenschaftlichen Klasse 4: 47–53.

    Google Scholar 

  • Krystyn, L. 1980. Stratigraphy of the Hallstatt region. In Second European Conodont Symposium (ECOS II), Guidebook, Abstracts, ed. H.P. Schönlaub, 69–98. Abhandlungen der Geologischen Bundesanstalt 35.

  • Krystyn, L. 1982. Obertriassische Ammonoideen aus dem zentralnepalesischen Himalaya (Gebiet vom Jomsom). Abhandlungen der Geologischen Bundesanstalt 36: 1–63.

    Google Scholar 

  • Krystyn, L., and Y. Gallet. 2002. Towards a Tethyan Carnian-Norian boundary GSSP. Albertiana 27: 12–19.

    Google Scholar 

  • Krystyn, L., Y. Gallet, J. Besse, and J. Marcoux. 2002. Integrated Upper Carnian to Lower Norian biochronology and implications for the Upper Triassic magnetic polarity time scale. Earth and Planetary Science Letters 203: 343–351.

    Article  Google Scholar 

  • Levera, M. 2012. The halobiids from the Norian GSSP candidate section of Pizzo Mondello (Western Sicily, Italy): Systematics and correlations. Rivista Italiana di Paleontologia e Stratigrafia 18: 3–45.

    Google Scholar 

  • Lopes, V.M., A.R. Lopes, P. Costa, and R. Rosa. 2013. Cephalopods as vectors of harmful algal bloom toxins in marine food webs. Marine Drugs 11: 3381–3409.

    Article  Google Scholar 

  • Lucas, S.G., N.J. Silberling, J.F. Jenks, J.A. Spielmann, and L.F. Rinehart. 2007. Third day: Upper Triassic and Lower Jurassic stratigraphy and biostratigraphy in western Nevada. In The Triassic of the American West, ed. S.G. Lucas and J.A. Spielmann, 23–30. New Mexico Museum of Natural History and Science Bulletin 40.

  • Lucas, S.G., L.H. Tanner, H.W. Kozur, R.E. Weems, and A.B. Heckert. 2012. The Late Triassic timescale: age and correlation of the Carnian-Norian boundary. Earth-Science Reviews 114: 1–18.

    Article  Google Scholar 

  • Martindale, R.C., D.J. Bottjer, and F.A. Corsetti. 2012. Platy coral patch reefs from eastern Panthalassa (Nevada, USA): Unique reef construction in the Late Triassic. Palaeogeography, Palaeoclimatology, Palaeoecology 313–314: 41–58.

    Article  Google Scholar 

  • Massare, J.A., and J.M. Callaway. 1988. Live birth in ichthyosaurs: Evidence and implications. Journal of Vertebrate Paleontology 8: 21A.

    Google Scholar 

  • Mazza, M., S. Furin, C. Spötl, and M. Rigo. 2010. Generic turnovers of Carnian/Norian conodonts: climatic control of competition?: Palaeogeography. Palaeoclimatology, Palaeoecology 290: 120–137.

    Article  Google Scholar 

  • Mazza, M., M. Rigo, and A. Nicora. 2011. A new Metapolygnathus platform conodont species and its implications for Upper Carnian global correlations. Acta Palaeontologica Polonica 56: 121–131.

    Article  Google Scholar 

  • Mazza, M., M. Rigo, and M. Gullo. 2012. Taxonomy and biostratigraphic record of the Upper Triassic conodonts of the Pizzo Mondello section (western Sicily, Italy), GSSP candidate for the base of the Norian. Rivista Italiana di Paleontologia e Stratigrafia 118: 85–130.

    Google Scholar 

  • McGowan, C., and R. Motani. 1999. A reinterpretation of the Upper Triassic ichthyosaur Shonisaurus. Journal of Vertebrate Paleontology 19: 42–49.

    Article  Google Scholar 

  • McLearn, F.H. 1960. Amonoid Faunas of the Upper Triassic Pardonet Formation, Peace River Foothills, British Columbia. Geological Survey of Canada Memoirs 311: 1–118.

    Google Scholar 

  • McMenamin, M.A.S., and D.L. Schulte McMenamin. 2011. Triassic Kraken: the Berlin Ichthyosaur Death Assemblage Interpreted as a Giant Cephalopod Midden. Geological Society of America Abstracts with Programs 43(5): 310.

    Google Scholar 

  • McMenamin, M.A.S., and D.L. Schulte McMenamin. 2013. The kraken’s back: new evidence regarding possible cephalopod arrangement of ichthyosaur skeletons. Geological Society of America Abstracts with Programs 45(7): 900.

    Google Scholar 

  • McMenamin, M.A.S., L.P. Zapata, and M.C. Hussey. 2013. A Triassic giant amphipod from Nevada, USA. Journal of Crustacean Biology 33: 751–759.

    Article  Google Scholar 

  • McRoberts, C.A. 2007. The halobiid bivalve succession across a potential Carnian/Norian GSSP at Black Bear Ridge, Williston Lake, northeast British Columbia, Canada. Albertiana 36: 142–145.

    Google Scholar 

  • McRoberts, C.A. 2010. Biochronology of Triassic Bivalves. In The Triassic Timescale, ed. S. Lucas, 201–219. London: Geological Society of London, Special Publication 334.

  • McRoberts, C.A. 2011. Late Triassic Bivalvia (chiefly Halobidae and Monotidae) from the Pardonet Formation, Williston Lake area, northeastern British Columbia, Canada. Journal of Paleontology 85: 613–664.

    Article  Google Scholar 

  • McRoberts, C.A. and L. Krystyn. 2011. The FOD of Halobia austriaca at Black Bear Ridge (northeastern British Columbia) as the potential base-Norian GSSP. In Canadian Paleontology Conference, Proceedings n. 9, ed. J.W. Haggart and P.L. Smith, 38–39. Vancouver: University of British Columbia.

  • Merriam, J.C. 1908. Triassic Ichthyosauria, with special reference to the American forms. Memoirs of the University of California 1: 1–196.

    Google Scholar 

  • Mojsisovics, E.v. 1874. Über Die Triadischen Pelecypoden-Gattungen Daonella und Halobia. Abhandlungen der kaiserlich königlichen Geologischen Reichsanstalt 7: 1–35.

    Google Scholar 

  • Monteiro, A., and P.R. Costa. 2011. Distribution and selective elimination of paralytic shellfish toxins in different tissues of Octopus vulgaris. Harmful Algae 10: 732–737.

    Article  Google Scholar 

  • Muller, S.W., and H.G. Ferguson. 1936. Triassic and Lower Jurassic formations of west central Nevada. Geological Society of America Bulletin 47: 241–252.

    Article  Google Scholar 

  • Muller, S.W., and H.G. Ferguson. 1939. Mesozoic stratigraphy of the Hawthorne and Tonopah quadrangles, Nevada. Geological Society of America Bulletin 50: 1573–1624.

    Article  Google Scholar 

  • Muttoni, G., D.V. Kent, P. Di Stefano, M. Gullo, A. Nicora, J. Tait, and W. Lowrie. 2001. Magnetostratigraphy and biostratigraphy of the Carnian/Norian boundary interval from the Pizzo Mondello section (Sicani Mountains, Sicily). Palaeogeography, Palaeoclimatology, Palaeoecology 166: 383–399.

    Article  Google Scholar 

  • Muttoni, G., D.V. Kent, P.E. Olsen, P. Di Stefano, W. Lowrie, S.M. Bernasconi, and F.M. Hernandez. 2004. Tethyan magnetostratigraphy from Pizzo Mondello (Sicily) and correlation to the Late Triassic Newark astrochronological polarity time scale. Geological Society of America Bulletin 116: 1043–1058.

    Article  Google Scholar 

  • Muttoni, G., M. Mazza, D. Mosher, M.E. Katz, D.V. Kent, and M. Balini. 2014. A Middle-Late Triassic (Ladinian–Rhaetian) carbon and oxygen isotope record from the Tethyan Ocean. Palaeogeography, Palaeoclimatology, Palaeoecology 399: 246–259.

    Article  Google Scholar 

  • Nicora, A., M. Balini, A. Bellanca, A. Bertinelli, S.A. Bowring, P. Di Stefano, P. Dumitrica, C. Guaiumi, M. Gullo, A. Hungerbuehler, M. Levera, M. Mazza, C.A. McRoberts, G. Muttoni, N. Preto, and M. Rigo. 2007. The Carnian/Norian boundary interval at Pizzo Mondello (Sicani Mountains, Sicily) and its bearing for the definition of the GSSP of the Norian Stage. Albertiana 36: 102–115.

    Google Scholar 

  • Orchard, M.J. 1991. Late Triassic conodont biochronology and biostratigraphy of the Kunga Group, Queen Charlotte Islands, British Columbia. In Evolution and Hydrocarbon Potential of the Queen Charlotte Basin, British Columbia, ed. G.W. Woodsworth, 173–193. Geological Survey of Canada Paper 1990–10.

  • Orchard, M.J. 2007. A proposed Carnian-Norian boundary GSSP at Black Bear Ridge, northeast British Columbia, and a new conodont framework for the boundary interval. Albertiana 36: 130–141.

    Google Scholar 

  • Orchard, M.J. 2013. Five new genera of conodonts from the Carnian-Norian Boundary beds, northeast British Columbia, Canada. In The Triassic system, ed. L.H. Tanner, J.A., Spielman, and S.G. Lucas, 445–457. New Mexico Museum of Natural History and Science, Bulletin 61.

  • Orchard, M.J. and E.T. Tozer. 1997. Triassic conodont biochronology, its calibration with the ammonoid standard, and a biostratigraphic summary for the Western Canada Sedimentary Basin. In Triassic of Western Canada Basin, ed. T. Moslow, and J. Wittenberg, 675–692. Canadian Society of Petroleum Geologists Bulletin 45.

  • Orchard, M.J., J.P. Zonneveld, M.J. Johns, C.A. McRoberts, M.R. Sandy, E.T. Tozer, and G.G. Carrelli. 2001. Fossil succession and sequence stratigraphy of the Upper Triassic of the Black Bear Ridge, northeast British Columbia, a GSSP prospect for the Carnian-Norian boundary. Albertiana 25: 10–22.

    Google Scholar 

  • Pyenson, N.D., C.S. Gutstein, J.F. Parham, J.P. Le Roux, C.C. Chavarría, H. Little, A. Metallo, V. Rossi, A.M. Valenzuela-Toro, J. Velez-Juarbe, C.M. Santelli, D. Rubilar Rogers, M.A. Cozzuol, and M.E. Suárez. 2014. Repeated mass strandings of Miocene marine mammals from Atacama Region of Chile point to sudden death at sea. Proceedings of the Royal Society B: Biological Sciences 281(1781): 20133316.

    Article  Google Scholar 

  • Robertson, A., D. Stirling, C. Robillot, L. Llewellyn, and A. Negri. 2004. First report of saxitoxin in octopi. Toxicon 44: 765–771.

    Article  Google Scholar 

  • Roniewicz, E., and G.D. Stanley. 2013. Upper Triassic corals from Nevada, western North America, and the implications for paleoecology and paleogeography. Journal of Paleontology 87: 934–964.

    Article  Google Scholar 

  • Sander, P.M. 2000. Ichthyosauria: their diversity, distribution, and phylogeny. Paläontologische Zeitschrift 74: 1–35.

    Article  Google Scholar 

  • Sandy, M.R., and G.D. Stanley Jr. 1993. Late Triassic brachiopods from the Luning Formation, Nevada, and their palaeobiogeographic significance. Palaeontology 36: 439–480.

    Google Scholar 

  • Seilacher, A. 1970. Begriff und Bedeutung der Fossil-Lagerstätten. Neues Jahrbuch für Geologie und Paläontologie Monatshefte 1970: 34–39.

    Google Scholar 

  • Seilacher, A. and F. Westphal. 1971. Fossil-Lagerstätten. In Sedimentology of parts of Central Europe, Guidebook 8. Int. Sediment. Congr. 327–335, Heidelberg.

  • Seilacher, A., W.E. Reif, and F. Westphal, 1985. Sedimentological, ecological and temporal patterns of fossil Lagerstätten. In Extraordinary fossil biotas: their ecological and evolutionary significance, ed. H.B. Whittington and S. Conway Morris, 311, 5–23 Philosophical Transactions of the Royal Society of London B.

  • Silberling, N.J. 1959. Pre-Tertiary Stratigraphy and Upper Triassic Paleontology of the Union District Shoshone Mounatins, Nevada. U. S. Geological Survey Professional Paper 322: 1–67.  

    Google Scholar 

  • Silberling, N.J. 1991. Allochthonous terranes of western Nevada, current status. In Geology and ore deposits of the Great Basin, ed. G.L. Raines, 101–102. Reno: Geological Society of Nevada.

  • Silberling, N.J., and E.T. Tozer. 1968. Biostratigraphic Classification of the Marine Triassic in North America. Geological Society of America, Special Papers 110: 1–63.

    Article  Google Scholar 

  • Silberling, N.J., D.L. Jones, Jr. M.C. Blake, and D.G Howell. 1987. Lithotectonic terrane map of the western coterminous United States. U.S. Geological Survey Miscellaneous Field Studies Map MF-1874-C, scale 1:2,500,000.

  • Smith, J.P. 1927. Upper Triassic marine invertebrate faunas of North America.U.S. Geological Survey Professional Paper 141: 1–262.

    Google Scholar 

  • Speed, R.C., N.J. Silberling, M.W. Elison, K.M. Nichols, and W.S. Snyder. 1989. Early Mesozoic tectonics of the western Great Basin, Nevada. In 28th International Geological Congress, Field Trip Guidebook T122: 1–54. Washington D.C.: American Geophysical Union.

  • Stanley Jr, G.D. 1977. Triassic Coral Buildups of Western North America. Lawrence: University of Kansas.

    Google Scholar 

  • Stanley Jr, G.D. 1979. Paleoecology, Structure, and Distribution of Triassic Coral Buildups in Western North America. Lawrence: University of Kansas Paleontological Institute.

    Google Scholar 

  • Tozer, E.T. 1965. Upper Triassic ammonoid zones of the Peace River foothills, British Columbia, and their bearing on the classification of the Norian stage. Canadian Journal of Earth Sciences 2: 216–226.

    Article  Google Scholar 

  • Tozer, E.T. 1967. A standard for Triassic time. Geological Survey of Canada Bulletin 156: 1–103.

    Google Scholar 

  • Tozer, E.T. 1971. Triassic time and ammonoids: Problems and proposals. Canadian Journal of Earth Sciences 8: 989–1031.

    Article  Google Scholar 

  • Tozer, E.T. 1981a. Triassic Ammonoidea: Classification, Evolution and Relationship, with Permian and Jurassic Forms. In The Ammonoidea, ed. M.R. House and J.R. Senior, The Systematic Association special volume 18: 65–100. London: Acadamic Press.

  • Tozer, E.T. 1981b. Triassic Ammonoidea: Geographic and Stratigraphic Distribution. In The Ammonoidea, ed. M.R. House and J.R. Senior, The Systematic Association special volume 18: 397–432. London, New York: Acadamic Press.

  • Tozer, E.T. 1984. The Trias and its ammonoids: the evolution of a time scale. Geological Survey of Canada, Miscellaneous Report 35: 1–171.

    Google Scholar 

  • Tozer, E.T. 1994. Canadian Triassic ammonoid faunas. Geological Survey of Canada Bulletin 467: 1–663.

    Google Scholar 

  • Wilby, P.R., J.D. Hudson, R.G. Clements, and N.T.J.J. Hollingworth. 2004. Taphonomy and origin of an accumulate of soft-bodied cephalopods in the Oxford Clay Formation (Jurassic, England). Palaeontology 47: 1159–1180.

    Article  Google Scholar 

  • Williford, K.H., M.J. Orchard, J.P. Zonneveld, C.R. McRoberts, and T.W. Beatty. 2007. A record of stable organic carbon isotopes from the Carnian-Norian boundary section at Black Bear Ridge, Williston Lake, British Columbia, Canada. Albertiana 36: 146–148.

    Google Scholar 

  • Zakharov, Yu.D. 1997. Carnian and Norian Sirenitid ammonoids of the north-western circum-pacific and their role in the Late Triassic faunal succession. In Late Paleozoic and Early Mesozoic Circum-Pacific Events: Biostratigraphy, Tectonic and Ore Deposits of Primoryie, (Far East Russia), ed. A. Baud, I. Popova, J.M. Dickins, S. Lucas and Yu.D. Zakharov, 137–144. Mémoires de Géologie 30.

  • Zonneveld, J.P., and M.J. Orchard. 2002. Stratal relationships of the Upper Triassic Baldonnel Formation, Williston Lake, northeastern British Columbia. Geological Survey of Canada Current Research 2002-A8: 1–13.

    Google Scholar 

  • Zonneveld, J.P., T.W. Beatty, K.H. Williford, M.J. Orchard, and C.R. McRoberts. 2010. Stratigraphy and Sedimentology of the lower Black Bear Ridge section, British Columbia: candidate for the base-Norian GSSP. Stratigraphy 7: 61–82.

    Google Scholar 

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Acknowledgments

We kindly thank the Nevada Division of State Parks and Nevada State Museum personnel who were responsible for expediting approval of our special permit application for paleontologic work within BISP. Their efforts are greatly appreciated. Likewise, we are appreciative of the US Forest Service for allowing us to conduct reconnaissance work on Toyable National Forest land. The on-site assistance of BISP supervisor Jeff Morris and his staff during our very rainy October 2010 field trip was also greatly appreciated.

Ammonoid taxonomic problems were discussed with Leo Krystyn (University of Vienna, Austria), who is the acknowledged leading authority on Carnian-Norian ammonoid systematics and biostratigraphy. Many thanks to Hans Hagdorn (Ingelfingen) for the stimulating discussions on concept and definition of mass mortality and Fossil-Lagestatten. Spencer G. Lucas and Justin A. Spielmann (New Mexico Museum of Natural History and Science, Albuquerque, NM) kindly helped with registration of the published specimens in the catalogue of the NMMNHS. Warm thanks also to Alexander Lukeneder (Naturhistorisches Museum, Wien), Hans Hagdorn and Leo Krystyn who kindly translated the Abstract into German.

The manuscript has been notably improved by valuable and stimulating suggestions by Leo Krystyn and Spencer G. Lucas.

Technical support for the ammonoid part was provided by G. Chiodi (University of Milano). This work is a contribution to PRIN 2008 project “Stratigrafia integrata del Triassico Superiore: GSSP e sezioni ausiliarie in Italia” (P.I. M. Balini). Financial support for field works has been provided by PRIN 2008 grants to M. Balini (Milano Research Unit). The conodont analyses were undertaken in Vancouver as a contribution to the Geological Survey of Canada’s “Geoscience for Energy and Minerals” (GEM) program, with support from the Yukon Basins Project (M.J. Orchard).

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Correspondence to Marco Balini.

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N. J. Silberling: deceased.

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Balini, M., Jenks, J.F., Martin, R. et al. The Carnian/Norian boundary succession at Berlin-Ichthyosaur State Park (Upper Triassic, central Nevada, USA). Paläontol Z 89, 399–433 (2015). https://doi.org/10.1007/s12542-014-0244-2

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