Robinson, R. Genetica (1970) 41: 190. doi:10.1007/BF00958904
The black form of the leopard is shown to be inherited as an autosomal monogenic recessive to the spotted wild type. Pairings of black animals inter se have a significantly smaller litter size than other possible pairings. Two reasons are offered for this: (1) that black animals may be slightly more inbred than the spotted or (2) black animals may be more prone to pre-natal loss than the spotted. The significance of the latter possibility is briefly discussed in relation to the persistence of the spotted/black polymorphism for areas of South-east Asia.